I noticed in last year's Tour de France (the only bike race I watch) that quite a few riders ride with their hands wrapped around the tops of their brake hoods and their forearms parallel to the ground. Apparently this riding position is "the most aero", at least compared to riding in the drops because it reduces the frontal area of one's forearms. Assuming this to be true, I think the "hoods-in" adjustment makes that position more comfortable as one's hands can rest a little more naturally on the brake hoods.
Since I started this post, I've kept my levers angled in and just love it. I only wish I had done this decades ago