If you were to put the same wheelset on each bike, that would eliminate that variable. With a properly-tuned and properly-tensioned wheelset, just about ANY frameset feels more 'sprightly', 'responsive' or 'faster' than the best frameset on a junk wheelset. If you don't believe me, just TRY it!
Take the most 'sluggish' bike you have and re-tension the wheels properly. Then you do the exact same ride. Which feels faster or more responsive? You'll also notice the increase in speed and less effort is used to achieve that faster speed.