When I look at this picture, I see that the bike is almost completely dismounted: crank, fork, headset, rear dropouts... even the tires must be dismounted from the wheels to reduce the wheels diameter in order to fit into the 62" box.
I am pretty sure that a Moulton with separable frame will fit with less dismounting.
Moulton doesn't claim that because they do not consider to dismount the bike, only separate the frame and maybe remove the wheels.
Note that the New Series are separable into more smaller pieces without dismounting any component:
Without mudguards, if the wheels are removed from the fork and rear frame, it packs very small.
But yes, the price is very high !