I'm surprised you even got off the starting line with that combination.
I didn't see the crank gearing listed on both. But, you likely have slightly different gearing.
So, on the Trek, you have: (12, 13, 14, 15, 16, 17, 19, 21, 24, 27)
On the Specialized, you (11-12-13-14-16-18-20-22-25-28-32T)
Not bad ratios on the 32T cassette, although you are missing the 15T and 17T on the Specialized. So it will depend a bit on which gear you settle into whether that would be an issue.
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Interesting comments on the Gatorskins. I like the Gator Hardshells, but I'm not a speed demon either.
I've been meaning to build up some kind of a deep tubular wheelset with high performance tires. Perhaps sometime.